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"From Mayer and Poljakoff-Mayber (1963) and Woody Plant Seed Manual (Anonymous, 1948).

"From Mayer and Poljakoff-Mayber (1963) and Woody Plant Seed Manual (Anonymous, 1948).

In angiosperm seeds, carbohydrates are stored in the endosperm, cotyledons, or both. In gymnosperm seeds, the reserves are stored primarily in the megagametophyte and cotyledons. In seeds of apple, ash, and dogwood, carbohydrates are stored primarily in the endosperm. In ailanthus, some reserves are stored in the endosperm and some in the cotyledons. The embryonic cotyledons of box elder and black locust store large amounts of carbohydrates. Those of ailanthus, green ash, and flowering dogwood store only small amounts. Species with seeds lacking endosperm (e.g., beech and black locust) have cotyledons adapted for both storage and photosynthesis (Marshall and Kozlowski, 1977). In Brazil nut, which has very small cotyledons, the embryonic axis is the major storage organ (Bewley and Black, 1978).

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