other extreme are species with very large leaves clustered at the ends of stout branches and with unbranched or few branched trunks (Figs 1 A, B). We refer to this growth form as pachycaul, a term which includes the Modèle de Corner and Modèle de Schoute of Hallé and Oldeman (1970) or the lamp post form of D'Arcy (1973). All species of Grias and several species of Gustavia have this growth form.

The classification of Lecythidaceae into pachycaul and leptocaul species is an oversimplification. For example, studies of the architecture of tropical trees by Hallé and Oldeman (1970) have revealed that our pachycaul category corresponds to two of their models and that our leptocaul group is more complex, including at least five of their architectural types. A summary of the architectural types of Hallé and Oldeman (1970) found in our two categories is as follows:

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