Structure of the Cytochrome b6f Complex Comparison with the Cytochrome bc1 Complex

The 217kDa hetero-oligomeric cytochrome b6 f complex [1] provides the electronic connection between Photosystem II [2, 3] and Photosystem I [4, 5] (see also Chapters 2, 3 and 4). Electron transfer through the b6 f complex is coupled to proton translocation to the electrochemically positive side of the complex (the luminal side) and, with proton deposition from the water splitting reaction of PSII and the decrease of proton concentration on the stromal side by reduction of NADP+ to NADPH, generation of a transmembrane proton electrochemical potential gradient, A(i + , marked by the p (lumen) and n (stromal) sides of the complex in Figure 7.1a.

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