Structure and Function of Cyanobacterial Photosystem I

Norbert Krauß

2.1 Introduction 23

2.2 Structural Overview 25

2.3 The Protein Subunits 27

2.3.1 The Core Subunits PsaA and PsaB 27

2.3.2 The Stromal Ridge: PsaC, PsaD and PsaE 30

2.3.2.1 Subunit PsaC 31

2.3.2.2 Subunit PsaD 32

2.3.2.3 Subunit PsaE 32

2.3.3 The Small Membrane-Integral Subunits 33

2.3.3.1 Subunits PsaL, PsaI, PsaM 34

2.3.3.2 Subunits PsaF, PsaJ, PsaX 35

2.4 The Electron Transfer Chain 37

2.4.1 The First Pair of Chlorophylls 39

2.4.2 The Second and Third Pairs of Chlorophylls 41

2.4.3 The Pair of Phylloquinones 43

2.4.4 The First [4Fe4S] Cluster, FX 46

2.4.5 The Terminal [4Fe4S] Clusters, FA and FB 46

2.5 The Core Antenna System 47

2.5.1 The Antenna Chlorophylls 48

2.5.2 The Carotenoids 51

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