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100'

Total sugar x 0.5

Hamamelitol

Sucrose

Glucose + Fructose Clusianose

| Fig. 1.3.26. Frost protection in the eastern Alpine Primula clusiana Tausch (A). Hamamelose = 2-hydroxymethyl-D-ribitol (B), and its galactoside clusianose (C), accumulate irreversibly in the overwintering leaves during the cold season (D). These old leaves die subsequent to the formation of new leaves in the following vegetation period without the branched chain sugar derivatives, which are present in high amounts, having been removed for further use. The accumulation of hamamelose can also be induced by cooling the plant in summer. (Photo E. Beck)

CH2OH HOH2C— C—OH H— C—OH H— C—OH CH2OH

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