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Energy flow

Definition and estimation Bryophytes and lichens are major elements in the tundra communities described in Chapter 2 in terms of cover and also energy flow, mineral nutrient cycling and other dynamic aspects of polar ecosystems. The part played by bryophytes has been briefly reviewed (Longton, 1984). Here we shall explore the role of both groups in greater detail, beginning with the contribution of cryptogams to production and phytomass.

Problems in Determining Reproductive Allocation

When the principle of allocation was formulated, two kinds of resources were in focus, energy and time. This, in combination with the general interest in energy flow in plant and systems ecology at that time (e.g., Odum, 1961), made energy a natural choice of currency to focus on also when studying plant reproductive allocation.

Traditional understanding of Magnolia bark Koboku

In Kampo medicine, Magnolia bark is believed to be a drug that replenishes KI or Chi (meaning vital energy, the basic function for existence). The common belief is that this plant can strengthen a patient's life energy by activating circulation of vital energy and eliminating dampness. Other claimed medicinal properties of Magnolia include diuretic and antitussive effects. Magnolia bark is used for pain in the abdomen due to entrapped gas and a feeling of congestion in the chest. It is also used in psychological disorders. A summary of the traditional understanding of Magnolia bark is presented in Table 2.2.

Ethnobotany Ethnopharmacology and Mass Bioprospecting

Traditional medicines play an important role in the provision of health care in many developing countries. Their use is also significant in developed countries, increasing their commercial value. Several 'highprofile' cases of patenting of traditional medicines, without consent from or compensation to their holders, have further focused attention on their importance. Traditional medicine usually involves biological resources and the knowledge of local and indigenous peoples and or healers regarding their medicinal use thus, it is interlinked with biodiversity conservation and indigenous peoples' rights over their knowledge and resources.

The European Dimension

The north-western forest culture includes Britain, Ireland, Belgium, the Netherlands, Denmark, southern Sweden, north-east France and Iceland. The common factor in these areas is the loss of most of their forest cover over the last 3-4 thousand years, so that forests now occupy a small percentage of the land area. Many forests are now of plantation origin and are intensively used. In popular culture forests are sometimes seen as alien places, so that urban people are less spiritually connected to them than in the Northern zone, for example. These are also highly urbanized and industrialized countries with dense populations who need space for recreation, but where private land is not always available for public access.

Role of Computer Modeling

Computer models have an extremely significant role in ecology. In fact, because so much in ecology is so difficult to measure except for the smallest plot, models are common in every field of ecology. Models are highly useful for testing scenarios. What will happen to stream flow and fish populations if 50 percent of trees are cut in a watershed How will elk populations change if wolves are reintroduced to a particular area How will the introduction of small controlled fires affect the potential for larger, highly destructive fires How will forests respond if carbon dioxide levels double in the next one hundred years These are just some of the ways that ecological models are used. see also Biome Carbon Cycle Clements, Frederic Ecology, Energy Flow Ecology, Fire

The Structure of an Ecological Community

Energy flow in an ecological community must obey the laws of thermodynamics. These constraints affect the flow of energy and therefore the structure of an ecological community. Using the grazing food chain as an example, let's see how these laws affect the flow of energy at each trophic level. For the harvest of solar energy by plants in the production of plant tissues, the first law of thermodynamics requires that the amount of solar energy captured by the plants remain the same before and after the transformation the energy involved cannot be created nor destroyed. For every thousand calories of solar energy captured and transformed by plant, there remain a thousand calories afterward. The second law of thermodynamics, however, requires that the harvesting of solar energy cannot be 100 percent efficient only a portion of the solar energy transformed by the plant can be conserved in the production of plant tissue. Measurements on various plant communities show that the actual...

Plastidic Adenine Nucleotide Transporters of Prokaryotic Origin

ATP represents the universal energy currency of all living cells, and is mainly synthesized in mitochondria and, in plant cells, also in chloroplasts. Mitochondria export ATP generated by oxidative phosphorylation in strict counterexchange with ADP via ADP ATP carriers (AAC). These carriers belong to the large mitochondrial carrier family (MCF), which in Arabidopsis consists of 58 proteins (Haferkamp 2007 Picault et al. 2004). An ATP transport activity has been shown to be present also in the envelope membranes of chloroplasts (Heldt 1969) and nongreen plastids (Sch nemann et al. 1993) and, recently, in the thylakoid membranes of chloroplasts (Spetea et al. 2004). Within the past decade, the proteins responsible for the plastid ATP transport activities have been identified and their physiological function partly characterized. It turned out that these proteins belong to two different unrelated transporter families, namely, the MCF and the plastid NTTs, the first having eukaryotic, the...

Conclusion and Future Goals

Nutrient fluxes are essential components of the material and energy flow that sustains ecosystems. Nutrient-seagrass interactions can be viewed under, at least, three perspectives (i) nutrient dynamics in seagrass beds (how nutrients enter, how they are used and recycled, and how they leave these ecosystems), (ii) influences of nutrient abundance on seagrass beds (how changes in nutrient supply alter functional aspects of the plant and modify ecological interactions), and (iii) influence of seagrass beds on nutrient cycles (how seagrasses modify the general nutrient fluxes in coastal waters). This chapter has focused mainly on the first two, and it is clear that a good understanding of them requires a multilevel approach, encompassing perspectives ranging from physiological, and even molecular, to ecological, and even bio-geochemical. The effects of nutrient additions provide a good example of this hierarchical nature of seagrass-nutrient interactions, which includes from changes in...

The Evolution of Feedback DeExcitation

And is rather similar to those of cyanobacteria. When cyanobacteria were engulfed by a eukaryotic host and became chloroplasts, the need for a respiratory chain creating a proton gradient over the thylakoid membrane disappeared (since energy and reducing power in the dark could be provided by the eukaryotic host), and photosynthetic electron transport became spatially separated from respiration. As a consequence, the proton gradient across the thylakoid became an indicator of excitation pressure and sensing of the lumenal pH became a direct way for an organism to detect whether electron transport or biochemistry limits photosynthesis. If biochemistry limits photosynthesis, excitation energy flow through PS I and PS II will be retarded and two adequate measures that a cell should take in order to avoid photodamage may occur. One is to increase the levels of photoprotectants in a wide sense and secondly, the efficient size of the light harvesting antenna can be decreased. A low...

Summary and prospectus

The 14C method for field estimation of rates of photosynthesis has provided an invaluable database for studies of the marine carbon cycle. However, nearly 50 years after its introduction to the discipline of biological oceanography, the original limitations of this approach continue to stand in the way of progress towards a comprehensive understanding of carbon and energy flow. New approaches for the routine and accurate estimations of GPCP, NPCP, NCCP and total energy flux are needed. It is the tradition in ecology to use carbon as the basic unit for biomass inventories and fluxes a more systematic and coordinated study of associated major and trace bioelements is also needed, including careful analyses of ecological stoi-chiometries and controls thereof. Only then can we begin to unravel the complex biogeochemical linkages that we seek to understand.

Introduction and overall aims

Energy flow through ecosystems has feedbacks to all spatio-temporal (fractal) levels. The lower levels (smaller in space, localised, shorter in time) always exhibit higher dynamics than upper or higher fractal levels (more space, longer in time). Thus, for example, longer-wave processes, such as forest-generation cycles or the successional development of ecosystems, only appear static from a short-term viewpoint. From a functional viewpoint, however, they should be considered rather as phases, whose harmonic patterns are simply revealed over longer time periods than those of shorter-wave subsystems.

Succession in Communities

Self-generating or autogenic succession leads to changes in community structure and ecosystem function. In the late 1960s, Eugene Odum described this as an overall strategy for ecosystem development. Even though general patterns of change appear to emerge, exceptions sometimes occur. There are, however, tendencies toward increases in biodiversity as succession progresses with slight declines as systems mature. Similarly, complexity and structure increase as succession proceeds, and increased proportional amounts of energy flow are needed to support increasing living community biomass there can be a tightening of nutrient cycling as the systems age. Thirty years later, Odum (1997) updated his thoughts in light of extensive research stimulated by the original model. In addition to systemic changes such as these, there are also plant life cycle strategies such as high seed number production, aggressive seed dispersal, high sunlight preferences, and rapid growth amongst invasive species...

Ecosystem efficiency of landscapes

How completely a landscape dissipates its daily input of solar energy is a measure of the efficiency of the entirety of its ecosystem(s). The extent to which the energy income to a delimited land surface (e.g. a catchment, subcatchment, or the domain of aggregated DEUs) and over a given time period (e.g. a year, or the complete lifecycle of longer-lasting primary producers such as forest trees) is dissipated is a measure of the efficiency of the whole ecosystem. However, ecosystem efficiency, too, has inherent limits. If the energy pulse of a certain frequency were fully damped within the given boundaries of a system (suppressing all spatio-temporal variability in energy flow and material supply, or matter budget, across the given surface), then matter-cycling would be perfect and loss-free, and the efficiency of the system would equal unity. The second law of thermodynamics, however, precludes this limit ever being reached (perpetual motion machines do not exist) and a certain degree...

Other effects of bryophytes and lichens

In addition to having a major impact on energy flow and nutrient cycling in polar ecosystems, bryophytes and lichens also interact with other organisms in a variety of different ways. For example, they provide a favourable habitat for invertebrates (Tilbrook, 1970), and lichens yield

Introduction and General Considerations

In some genera within the Gramineae, as in some sugarcane varieties, quite substantial BNF does occur without any inoculation (Boddey 1995). In addition, there is also strong evidence from other gramineous plants like Pas-palum notatum cv. batatais, the smooth cord grass (Spartina alterniflora), Kallar grass (Leptochloa fusca) in Pakistan, and dune grass (Ammophila arenaria and Elymus mollis) from Oregon (Dalton et al. 2004) growing in nitrogen-limited soils, that endophytic BNF can contribute substantially to the nitrogen need of Gramineae. In the case of Paspalum notatum, the 15N isotope-dilution method showed that 10 of the total N accumulated in the grass originated from BNF (Boddey et al. 1983). A high diversity of dia-zotrophic bacteria have been isolated over the years from rhizosphere soil and the root surface (rhizoplane) of these plants, but none of these diazotrophic bacteria could be found to contribute considerable amounts of fixed nitrogen to its host plant, thus...

Herbal Treatments For Adrenocortical Dysfunction

To compensate for increased stressors, many individuals have turned to ergogenic or energy enhancement substances. These herbal and nutritional supplements are thought to have some type of ergogenic activity and are among the best-selling natural products in nutrition stores, with a financial impact in the 2- 3 billion per year range. Although there is a body of scientific literature on a variety of natural ergogenic substances such as pyruvate, creatine, ephedra (ma huang Ephedra sinica), ginseng (Panax spp.), and guarana (Paullinia cupana) using animal models, there are few well-designed human clinical trials. This lack of legitimate research and the high over-the-counter use of natural products suggest an urgent need to conduct studies on the long-term effectiveness and safety of these natural ergogenic aids. Natural products (such as phytopharmacologic agents), which appear to enhance performance capacity (as demonstrated in animal and human studies), include such nutrients as...

The cardiovascular system

Most herbalists will know that many plants in this group (several formerly used as arrow poisons1 or even in ordeals) are very toxic and will avoid using them. A few healers, however, may, in view of the fact that they are also emetics, purgatives or diuretics, be tempted to make use of them. But these plants should be employed only after complete extraction and with very exact dosages of the active constituents, and then only by physicians in possession of a full clinical diagnosis. In this, these plants differ from many others, which may be given as a decoction, an infusion or in dried or powder form.

What Distinguishes Plants From Medicinal Plants

Ayurveda, the ancient healing system flourished in India in the Vedic era. The classical texts of Ayurveda, Charaka Samhita and Sushruta Samhita were written around 1000 B.C. The Ayurvedic Materia Medica includes 600 medicinal plants along with therapeutics. Herbs like turmeric, fenugreek, ginger, garlic and holy basil are integral parts of Ayurvedic formulations. The formulations incorporate a single herb or more than two herbs (polyherbal formulations). and is a popular science in western countries. Some of the medicinal herbs mentioned in Chinese medicine are common to Ayurveda. Traditional Chinese medicine favors the use of medicinal herbs in their natural form rather than by extraction. Herbal drugs have a different history in Europe and America and have produced healers like Culpeper. The use of tinctures in homeopathy is based on medicinal herbs. Ancient healers, who developed formulations based on medicinal herbs, were probably not aware of the chemical composition of the...

Role for Physicians in Ethnopharmacology and Drug Discovery

Ethnopharmacology investigations classically involved traditional healers, botanists, anthropologists, chemists and pharmacologists. The role of some groups of researchers but not of physicians has been highlighted and well defined in ethno pharmacological investigations. Historical data has shown that the discovery of several important modern drugs of herbal origin owe to the medical knowledge and clinical expertise of physicians. Current trends indicate a negligible role of physicians in ethnopharmacological studies. (1) Ethnopharmacological field work which involves interviewing healers interpreting traditional terminologies into their modern counterparts examining patients who are consuming herbal remedies and identifying the disease for which an herbal remedy is used. (5) Work in collaboration with local healers to strengthen the traditional system of medicine in a community.

Understanding the Impact of Soil Organic Matter on Suppressiveness

The main energy channels within the detritus food web are either bacterial or fungal, and soil ecologists suggest that the dominant channel in natural systems is largely determined by litter quality and the environment (Bardgett 2005 Wardle 2005). However, in agroecosystems, soil management practices also determine whether energy flow occurs rapidly through bacterial channels or more slowly through fungal channels. Tillage and nitrogen fertilisation practices have particularly profound effects on the soil food web, stimulating bacterial activity to such an extent that bacterial rather than fungal decomposers predominate in many farming systems. Changes in the proportions of bacteria to fungi and their flow-on effects to other components of the soil food web, together with the detrimental effects of tillage (Wardle 1995) and nitrogen (Tenuta and Ferris 2004) on some predators may explain why soils tend to lose their natural suppressiveness once they are cropped. These issues need to be...

Other Species Of Artemisia Used In Traditional Chinese Medicine

More than ten Artemisia species are used in TCM for certain gynaecological problems. According to the theory of TCM, a number of conditions, such as amenorrhea, menstrual pain and prolonged menstrual bleeding, are usually related to Qi (vital energy) and blood deficiency, Qi stagnation or blood stasis due to cold. Thus those herbs with the acrid and warm property can be used to treat such ailments with a good clinical response and various species of Artemisia which have the same property are often the principal ingredients of TCM formulae for the illnesses mentioned above (Table 1).

Early traditional medicine

The if a may say that he has discovered a plant possessing a spirit stronger than the disease spirit, and he and his patients believe that the power of this spirit, or the soul of the medicine, is not manifested before the healer has spoken some magic words or has chanted an incantation over the plant. Before doing so the if a himself may appeal for advice to gods or worship idols which in Yoruba country (Western Nigeria) are often small carved figures of a man or woman and sometimes also of animals. The patients in turn should not only take or apply the medicine but also appeal to and make offerings to communal and household gods, which may also be carved statues or other objects blessed by the local priests in ceremonies that generally last for days. In a school in Badagry (Nigeria, near the frontier with Dahomey) there were in a dark corner places of sacrifice consisting of cones of clay with an irregular shiny surface. They were streaked in white, turquoise, yellow and brown by...

Herbal Medicine Today

Today, traditional herbalist healers continue to use knowledge passed down for generations. Some ethnobotanists are studying with traditional healers to save such knowledge before it disappears. Arvigo, Rosita, Nadine Epstein, Marilyn Yaquinto, and Michael Balick. Sastun My Apprenticeship with a Maya Healer. San Francisco HarperSanFrancisco, 1994.

Better Sex Naturally

Traditional healers and physicians around the world have long known that there are many natural medicines that can enhance sexual desire and function in both men and women. The Chinese have used ginseng for thousands of years as a tonic and to stimulate desire and enhance endurance. Similarly, the herb known as Indian ginseng, ashwagandha (Withania somniferum), has been used to promote potency for centuries. Throughout the world, one finds an abundance of long-revered aphrodisiacs that have now gained increased acceptance as a result of clinical trials. However, from a natural medicine perspective, the best approach to enhancing bodily functions including sexuality is to support the underlying health of the body as a whole. It is true that the better our patients feel, the greater will be their ability to enjoy sexual satisfaction.

Folk Recipes

Just like allopathic and homeopathic medicine, the traditional herbal system uses a special combination of plants to treat diseases. Different plants have different chemicals that can have different results on different organisms. The proper quantity and quality of these plants must be consumed in order to get positive results otherwise, either no results or some mishap can occur. That is why local people are always in search of authentic recipes. Today a lot of people are using the traditional system for example, in China, traditional Chinese medicine is relied upon for nontoxicity, and most Chinese people avoid allopathic or other systems. In the Lesser Himalayas, local inhabitants use medicinal plants in different ways, s in the forms of decoctions, extracts, powders, pastes, or juices. The mode of application for these plants is topical as well as oral. Different recipes used by the indigenous people of the Lesser Himalayas have been developed from time immemorial through...

Ethnobotany

For the history of the uses of this plant in the period before and after the conquest we have to rely on some colonial codices. The best known ones are the Codex Cruz Badiano and the Codex Florentino. The first is a herbal written in Nahuatl by the Aztec healer Martin de la Cruz from Tezcoco, who was at the Colegio de Santa Cruz in Tlatelolco. It was translated into Latin by Juan Badiano and given to the King of Spain Carlos I in 1552. It was written rather hastily and has numerous colour illustrations of medicinal plants. There have been several attempts to identify plants from this herbal (Viesca Trevino, 1992 Valdes et al., 1992 Pineda, 1992) and most of the identifications seem to be botanically sound. The major problem with this source is that by this time the European conquest of Mexico-Tenochtitlan 30 years previously had already had an impact. In addition the Nahuatl author attempted to show European sophistication in his work (Ortiz de Montellano 1990).

Food Webs

In the 1950s the idea of the food web began to emerge in ecosystem ecology. Food webs and the related topics of trophic levels and energy flow are some of the most critical ecological concepts because they illustrate the connections between organisms that are required to maintain healthy ecosystems. Energy flow refers to the way that energy is transformed through a food chain containing a series of levels, including plants, consumers, predators, and decomposers. Each step in the food chain is called a trophic level (from the Greek word trophikos, meaning nutrition ). Primary producers (plants, algae, and photosynthetic microbes) are the base of food chains and are the lowest trophic level. They transform energy from the Sun into sugars. Primary producers thus make their own food and are called autotrophs all other organisms ultimately use the energy produced by autotrophs and are called heterotrophs. At the next trophic level, primary consumers (herbivores) eat some of the sugars...

Concluding remarks

The EO of L. angustifolia (Lamiaceae) has been used since ancient times on account of its healing properties in the treatment of various diseases, ailments, disorders and discomforts as well as in cosmetic formulations and in perfumery. By means of these properties lavender flowers as well as their oil are even accepted in various pharmacopoeas (see also Wichtl, 1997). Aromatherapy as well as aromachology make use of this EO to a great extent and also nearly all alternative healing treatments (scientific and esoteric and unscientific ones) know of its beneficial activities. The characteristic woody-herbaceous odour with green-floral notes creates a sort of 'well' feeling in the user by which it can be used as a mood tonic, antidepressant and psychological remedy, for example, against shock of injury (Worwood, 1994). In that way the aromachological usage of this EO is equally important as its use in (scientific) aromatherapy (see Buchbauer et al., 1994 Buchbauer, 1998). As can be seen...

Ecosystems

The beauty of ecosystem models is that they can be quantified. This allows them to be analyzed mathematically on computers and ultimately, if the models are based on real, natural behaviors, they can be used to predict the future of ecosystems. The rapidly developing field of general systems theory can be applied to ecosystems resulting in insights about how they function. These tools also allow ecologists to make predictions about the behavior of ecosystems when disturbed, stressed, or altered by evolutionary time, questions that society is finding pressing with increasing pollution, global warming, and other environmental threats. see also Agricultural Ecosystems Aquatic Ecosystems Biome Coastal Ecosystems Ecology, Energy Flow Ecology, Fire Ecology, History of Plant Community Processes.

Odum Eugene

Odum also wrote several other works while teaching and doing research at the University of Georgia. His work was funded by the Atomic Energy Commission, an institution that funded much early ecological research. He became a leading authority on ecosystem studies, defending the new discipline against its critics, and he also served as chair of a section of the International Biological Program. His leadership in the program helped guide research into landscape ecosystems, studying terrestrial and marine areas and the human influences on them. Remaining active into his late eighties by the turn of the twenty-first century, Eugene Odum still worked to promote the study of ecosystems. He has done much to encourage environmental study around the world, and especially where he works in Georgia. see also Ecology Ecology, Energy Flow Ecology, History of Ecosystem Warming, Johannes.

ATP Production

In plants essentially all electron flow from water follows the pathway shown in Figure 3, at least up to ferredoxin. However, once an electron reaches ferredoxin the electron pathway becomes branched, enabling a fraction of the redox free energy to enter other pathways, including cycling through the Photosystem I reaction center. Photosystem I cyclic electron transport provides additional energy for ATP production, which allows plants to adjust the energy flow according to their metabolic needs.

Linker Proteins

With the crystallographic visualizations of the PC and PEC trimeric hexameric structures, it became clear that if the linkers are present in the disk cavity, they will come into contact with the chromophores that jut out into this solvent accessible space (for instance, the PPCB84 in PC, PEC, and APC). Since the chromo-phore environment has a very strong effect on its spectral characteristics, the presence of different LPs could potentially modify the functionality of these similar, or even identical, chromophores. By specifically altering the environment of certain chromophores, the LPs could induce directed energy flow down the rods and into the core, and from there into the reaction center. Thus, a second role for LPs was proposed. Indeed, biophysical measurements made on isolated PBPs show a significant shift in their absorption spectra in the presence or absence of LPs '41 - No phycobilisome has yet been isolated that does not contain LPs but, mutagenesis that causes the lack of...

Plant Prospecting

Botanists follow either random or targeted approaches when choosing plants for pharmacological studies and drug discovery. The random prospecting strategy is to gather all of the available vegetation in an area supporting rich biological diversity. The more focused methods are taxo-nomic, ecological, and ethnobotanical. The taxonomic method emphasizes the collection of close relatives of plants already known to produce useful compounds for medicine or other uses. The ecological approach focuses on plants that offer certain clues promissory of activity, such as plants free from herbivore predation, which imply the presence of chemical defenses. Finally, ethnobotanical prospecting is done by interviewing native healers who have knowledge of the local plant's medicinal properties.

Nutrient cycling

These effects, and the youthfulness of polar ecosystems, result in available nitrogen and phosphorus being limiting in many tundra soils. Indeed, low rates of nutrient cycling could impose restrictions on productivity as severe as those attributable to direct effects of temperature on metabolism in the producer organisms (Chapin, 1983 Heal et al., 1981). The relative importance of these factors is not clear, as nutrient cycling has so far been less intensively studied than energy flow (Dowding et al., 1981).

The nervous system

I still consider it important to classify the plants by their clinical indications, in spite of the fact that the greatest amount of research is still to come, to keep in mind the aim, and to see rapidly and more clearly which of the far too numerous utilizations of each plant by local healers appear the most important and are confirmed by scientific observations. The enormous treasure of natural remedies should not be used in a haphazard way. A simple extraction might in some cases enable a healer to increase or isolate a fraction with a certain action or to eliminate a toxic constituent. The experience with the same plants in similar climates even in different continents should not be ignored (although the amount or quality of the constituents can vary and should be checked).

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