multiple genes for salt tolerance whose impact varies with stress duration, leaf age, and leaf type. Lang et al. (2001) reported a micro-satellite marker, RM223 on rice chromosome 8, associated with salt tolerance at both vegetative and reproductive stages. Recently, Alam et al. (2011) identified salinity-related QTLs at seedling stage using SES scores and identified Saltol as well as non-Saltol related QTLs at seedling-stage in Pokkali, which may assist in QTL pyramiding and marker-assisted breeding programs. A study by Lin et al. (2004) employing the tolerant indica landrace Nonabokra with the susceptible japonica Koshihikari, identified several large-effect QTLs, including the SKC1 QTL and a QTL for shoot Na+ concentration. While the salt-tolerant landraces Pokkali and Nonabokra were routinely used in the past for breeding, the level of tolerance attained by new lines is always lower than traditional donors (Gregorio et al. 2002), and the existing tolerant varieties seem to be superior in only few traits associated with tolerance. The QTL SKC1, originally detected by its effect on K+ concentration, was cloned by map-based cloning and was

Table 4.3 Molecular markers associated with quantitative trait loci for salt tolerance in barley

Molecular markers




Chr 1H


Seedling salt tolerance

Steptoe X Morex

Mano and Takeda (1997)


Seedling salt tolerance

Steptoe X Morex

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